Weird layer issues in my print

[lolad]

I've got these weird gaps appearing in my layers, as well as gaps in the top layer. Any idea why? (Image of calibration cube print in attachments)Ender 3 V2 Neo, PrusaSlicer, PLA. I've printed fine on this for several months and just replaced the hotend internals after wrecking the previous one. Running at 220 degrees - I have tried lower as I usually print slightly lower but this was the best attempt so far (bed 60C, and I've tuned PID).I've calibrated esteps. I don't think it's underextrusion as I've tried running a cube with a higher extrusion multiplier (1.2) which doesn't fix the problem and causes typical overextrusion symptoms.I've got 5 top layers in the slicer and fairly default settings as I reset the profile to default to see if the problem was caused by me setting a weird slicer setting.Filament is not tangled, nozzle is not clogged (as far as I am aware), bed is levelled with ABL and first layer is fine and solid, unlike the top layer. The extruder tension is good too as the marks on the filament show a firm grip but not grinding.Infill is at 10% adaptive cubic - which usually seems to work fine. I don't suspect filament hydration as it's only a few months old and it's PLA (eryone). I've kept it on the shelf and it's not been too humid so I wouldn't expect it to be wet - I've tried this with polyterra pla and I get similar results. I don't see bubbles or anything when it's extruding.I'd really appreciate any ideas as I seem to have tried just about everything. Thanks!