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It looks like you have Z wobble/backlash. Are your Z rods constrained at the top and bottom? Do they sway as the Z goes up and down?

You might look into using this...

I used a spring clamp like this

3228 to pull the Y motor back to tighten the Y belt, but I can't do the same for the X motor.

I jokingly thought up a way to communicate exactly how tight the...

Don't. I got a couple of these http://e3d-online.com/Mechanical/GT2-Belt-Tensioning-Spring, and as far as I could tell, they didn't improve prints. Rather, they, being springs and bouncy, ended up...

Any advice for getting the belts to the "correct" tension? I've been struggling with it lately, and circular objects I've printed are coming out not-so-circular. I keep hearing how belts shouldn't be...

Try adjusting your bridging speed. In Slic3r, I bridge at 10mm/sec. You may also need a fan to blow on the filament as it's extruded while bridging to help cool it off quicker.